Vacancies

The PERICO research programme aims to uncover how a central metabolic organelle, the peroxisome, participates in controlling cellular metabolism. Metabolism and metabolic control are emerging as important frontiers in physiology. This is because, in our rapidly changing world, we face the effects of both metabolic extremes – from obesity on the one hand to malnutrition on the other. Learning how to control energy metabolism is therefore one of the most pressing challenges of the 21stcentury.

Peroxisomes are key metabolic organelles, which must communicate and interact extensively with their environment to exchange metabolites and coordinate cellular responses. Membrane contact sites (MCS), where membranes of two organelles are physically tethered to enable rapid transfer of small molecules, enable organelle communication and are crucial for coordination of cellular functions and hence human health. Research on organelle interactions and communication is a challenging, upcoming field in current cell biology.

PERICO will exploit recent developments in high-throughput and genome-wide screening technologies, combine these with modern molecular cell biology and systems biology and ultimately translate the data into new leads for drug discovery and therapy.

15 Early Stage Researchers

The training schemes include specific research projects, secondments to partner organizations, a wide range of dedicated courses, and workshops organized by the academic and industrial partners of the Network. The scientific training programme will be complemented with training in managerial soft skills.

The positions for Early Stage Researchers are available for candidates with a research experience ≤ 4 years (counted from the diploma that gives the rights to embark in a doctoral degree).

Candidates must not have resided or carried out their main activity (work, studies, etc) in the country of their host organisation for more than 12 months in the 3 years immediately prior to recruitment (short stays, such as holidays, are not taken into account).

Requirements

Candidates should have:

  • exceptional academic performance, including qualifications, prizes
  • subject specific skills and expertise (see project descriptions)
  • communication, presentation skills and team working abilities
  • competence in written and spoken English.

Length of appointment: 3 years
Type of contract: temporary
Application deadline: 28/10/2018
Starting date: April 1, 2019

Applicants can apply for multiple positions.

More information on the project

Descriptions of the individual PhD projects

Reference: ESR1-NL

A PhD position is available at the Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, The Netherlands.
Project title: Peroxisome-vacuole membrane contact sites (MCS) in yeast
Description:

  • Comparison of wild-type and peroxisome-deficient yeast cells by metabolomics/lipidomics to identify how peroxisome deficiency affects cell metabolism and signalling networks.
  • To identify novel contact site proteins using proteomic approaches and bioinformatics search for homologues
  • Validation of selected, putative yeast contact site proteins using advanced microscopy techniques including correlative light and electron microscopy
  • Detailed functional analysis of selected novel vacuolar contact site proteins (peroxisome biogenesis, dynamics and metabolism) in mutant strains using biochemical, microscopy and “omics’ approaches.

Location: Groningen, The Netherlands
Supervisor: Prof. Dr. I.J. van der Klei (i.j.van.der.kleiATrug.nl)
Co-supervisors: Prof. Dr. R. Erdmann & Dr. M. Danielsen (MS-Omics ApS)
Planned secondments: Ruhr University Bochum, University of Freiburg, MS-Omics ApS
Required subject specific skills and expertise: a master’s degree in molecular life sciences (molecular cell biology, molecular genetics, biochemistry). Experience in yeast research is advantageous, but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR 1

Reference: ESR2-NL

A PhD position is available at the Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, The Netherlands.
Project title: Yeast peroxisomal transporters and pore forming proteins
Description:

  • Identification of novel transporters by proteomic analysis of peroxisomal membranes.
  • Library construction and high-throughput fluorescence microscopy screens to identify peroxisomal transporters.
  • Identification of potential novel yeast peroxisomal transporters using bioinformatics.
  • Validation and functional analysis of selected candidates.

Location: Groningen, The Netherlands
Supervisor: Prof. Dr. I.J. van der Klei (i.j.van.der.kleiATrug.nl)
Co-supervisors: Mr. J. Wehmeijer, Lambert Instruments & Prof. Dr. M. Schuldiner, Weizmann Institute of Science
Planned secondments: MS-Omics ApS, Weizmann Institute of Science, Lambert Instruments
Required subject specific skills and expertise: a master’s degree in molecular life sciences (molecular cell biology, molecular genetics, biochemistry). Experience in yeast research is advantageous, but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR 2

Reference: ESR3-NL

A PhD position is available at the University Medical Centre Groningen, Department Paediatrics, Centre for Liver, Digestive and Metabolic Diseases, The Netherlands.
Project title: Translational models for peroxisomal dysfunction in genetic disease and malnutrition
Description:
In this ‘Systems Medicine’ project we will focus on the construction of computational models to understand how severe acute malnutrition (SAM) leads to sustained dysfunction of peroxisomal and mitochondrial metabolism. A challenge for such an approach is to bridge the gap between computational models and the in vivo metabolism in patient tissues. To this end the PhD student will also develop in vitro disease models, such as liver organoids and iPS-derived liver cell culture. Tasks include:

  • Establish an in vitro model for SAM (liver iPS cells and/or organoids in medium deprived of amino acids).
  • Biochemical characterization of the in vitro malnutrition model, focusing on fluxomics and proteomics.
  • Construct a computational model of the interplay between peroxisomes and mitochondria in lipid metabolism.
  • Model-informed interventions to rescue peroxisomal-mitochondrial function.

Location: Groningen, The Netherlands
Supervisor: Prof. Dr. B.M. Bakker (b.m.bakker01AT umcg.nl)
Co-supervisors: Prof. Dr. V. Martins Dos Santos, Lifeglimmer GmbH & Dr. R. Bandsma, The Hospital for Sick Children
Planned secondments: The Hospital for Sick Children, KU Leuven, Lifeglimmer GmbH
Required subject specific skills and expertise: A masters’ degree in life sciences, biophysics, biochemistry, or a related field, with clear affinity for metabolism and mathematics.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR3

Reference: ESR4-NL

A PhD position is available at the Laboratory Genetic Metabolic Diseases, Academic Medical Centre at the University of Amsterdam, The Netherlands.
Project title: Human peroxisomal metabolite/cofactor transporter proteins
Description:

  • To identify potential human transporter proteins by proteomic analysis of purified human peroxisomal membranes.
  • Identification of promising candidate proteins using bioinformatics.
  • Validation of selected candidates and effect on metabolism (CRISPR, fluorescence microscopy and electron microscopy, cell fractionation, metabolomics);
  • Functional analysis of novel and selected known transporter proteins in vivo to determine role in human metabolism (CRISPR, metabolite sensors, metabolomics);
  • Identification and characterization of novel or previously described human disorders caused by defects in genes encoding peroxisomal metabolite transporter proteins (whole exome sequencing data and databases/bioinformatics);
  • Search for compounds that can rescue or bypass human transporter defects in cells.

Location: Amsterdam, The Netherlands
Supervisor: Prof. Dr. H.R. Waterham (h.r.waterhamATamc.uva.nl)
Co-supervisors: Dr. D. Hoepfner, Novartis Pharma AG & Prof. Dr. M. Fransen, KU Leuven
Planned secondments: Novartis Pharma AG, KU Leuven, University of Exeter
Required subject specific skills and expertise: a master’s degree in life sciences/biology (molecular cell biology, biochemistry, biomedicine). Experience in mammalian cell culture and fluorescence microscopy is advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR4

Reference: ESR5-NL

A PhD position is available at Lambert Instruments, Groningen, The Netherlands
Project title: Autonomous High-Throughput accurate FLIM with subcellular resolution
Description:
Development and optimization of automated data mining from multi-well microscopy plates.
The goal is to improve on current intensity-based methods for detection of the presence of biological phenomena in microscopic images by applying deep-learning for morphological analysis in detection and classification. Additional discriminating dimensions are created by integrating Fluorescence Lifetime Imaging Microscopy (FLIM) measurements into the dataset. The end point is a fully automated processing workflow to detect subtle changes in organelle morphology. The research involves improvement of a FLIM protocol for speed and accuracy, optimize modulation and illumination parameters, develop noise reduction algorithms. By using active feedback loops for real-time measurement adjustment, the harvest of target patterns will be optimized using reinforcement learning or other control algorithms for digital microscopes.

Location: Groningen, The Netherlands
Supervisor: Prof. Dr. L.R.B. Schomaker (l.r.b.schomakerATrug.nl); Mr. J. Wehmeijer (jeroenATlambertinstruments.com)
Co-supervisors: Prof. Dr. I.J. van der Klei, University of Groningen & Prof. Dr. M. Fransen, KU Leuven
Planned secondments: KU Leuven, University of Groningen
Required subject specific skills and expertise: a master’s degree in computer science or artificial intelligence with specialisation in image processing and (deep) machine learning. Experience with biological imaging techniques is advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR5

Reference: ESR6-DE

A PhD position is available at the Department of Systems Biochemistry, Ruhr-University of Bochum, Bochum, Germany
Project title: Novel transporters of Trypanosoma glycosomes
Description:

  • Identification of Trypanosoma transporters using proteomics approaches (proteomics of purified organelle membranes and isolated Pex19 complexes) and in silico screening of homologs of known transporters.
  • Validation of the suitability of selected transporters as potential novel drug targets (RNAi or CRISPR to determine essentiality for trypanosome survival).
  • Disruption of glycosomal targeting of transporters by block of interaction between Pex19 and sorting signal of membrane proteins (mPTS) as novel drug target.
  • Identification of mPTS in transporters, establishment of the in vitro interaction assay, drug/compound library screening (in cooperation with Novartis Pharma AG).

Location: Bochum, Germany
Supervisor: Prof. Dr. R. Erdmann (ralf.erdmannATrub.de)
Co-supervisor: Dr. D. Hoepfner, Novartis Pharma AG & Prof. Dr. H. R. Waterham, Academic Medical Centre at the University of Amsterdam
Planned secondments: University of Freiburg, Novartis Pharma AG, Academic Medical Centre at the University of Amsterdam
Required subject specific skills and expertise: a master’s degree in life sciences/biology (molecular cell biology, biochemistry, biomedicine). Experience in cell culture and fluorescence microscopy is advantageous but not mandatory.

Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR6

Reference: ESR7-DE

A PhD position is available at the Department of Systems Biochemistry, Ruhr-University of Bochum, Bochum, Germany
Project title: Glycosomal membrane contact sites in trypanosomes
Description:

  • Characterisation of the ATAD1/Msp1-derived membrane contact site (MCS) and identification of novel Trypanosoma MCS proteins using proteomic approaches (Affinity isolated protein complexes of ATAD1/Msp1 and Pex11) and by screening for trypanosomal homologues of newly identified MCS proteins.
  • Validation of the suitability of selected MCS proteins as potential drug targets (RNAi knockdown/CRISPR in Trypanosoma, metabolomics icw MS-omics, systems biology models (in cooperation with UMCG).
  • Screening of drug/compound library (in cooperation with Novartis Pharma AG) for specific inhibitors of Trypanosoma ATAD1/Msp1 ATPase activity.

Location: Bochum, Germany
Supervisor: Prof. Dr. R. Erdmann (ralf.erdmannATrub.de)
Co-supervisors: Dr. D. Hoepfner, Novartis Pharma AG & Prof. Dr. B. Warscheid, University of Freiburg
Planned secondments: University of Freiburg, Novartis Pharma AR, Oroboros Instruments
Required subject specific skills and expertise: a master’s degree in life sciences/biology (molecular cell biology, biochemistry, biomedicine). Experience in cell culture and fluorescence microscopy is advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR 7

Reference: ESR8-DE

A PhD position is available at the University of Freiburg, Department of Biochemistry & Functional Proteomics, Freiburg, Germany
Project title: Dynamics and regulation of ER-peroxisome contact sites
Description:

  • Mapping of in vivo phosphorylation sites in Pex30p-dependent ER-peroxisome contact sites (EPCONS) under different conditions in yeast using high-resolution mass spectrometry.
  • Identification of kinases mediating phosphorylation events and their preferred target sites to decipher the cellular signalling pathway(s) regulating EPCON formation.
  • Functional analysis of EPCONS using site mutants of selected proteins mimicking or preventing phosphorylation or targeting selected protein kinases and signalling pathways using conditional mutant strains or specific kinase inhibitors.
  • Characterizing human PEX30 and its role for the formation of EPCONS.

Location: Freiburg, Germany
Supervisor: Prof. Dr. B. Warscheid (bettina.warscheidATbiologie.uni-freiburg.de)
Co-supervisors: Prof. Dr. L.R.B. Schomaker, Lambert Instruments & Prof. Dr. I.J. van der Klei, University of Groningen
Planned secondments: University of Exeter, Lambert Instruments, University of Groningen
Required subject specific skills and expertise: a master’s degree in biology, biochemistry or a related discipline. Experience in proteomics/mass spectrometry and molecular genetics is advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR8

Reference: ESR9-DE

A PhD position is available at the University of Freiburg, Department of Biochemistry & Functional Proteomics, Freiburg, Germany
Project title: High-resolution proteomics of peroxisome-centred membrane contact site (MCS) and transporters
Description:

  • Definition of the peroxisomal membrane proteome of yeast, human/mammalian cells, and trypanosomes using high-resolution quantitative mass spectrometry (MS) to identify putative new transporters.
  • Screen for novel membrane contact site (MCS) proteins in yeast using artificial protein tethers, quantitative MS, and computational data analysis.
  • Interactomics and proximity proteomic studies of key proteins forming MCS in different organisms and/or under different growth conditions.
  • Global loss-of-function studies of peroxisome-deficient yeast cells and cells with non-functional MCS and transporters and analysis of the cellular response to the lack of peroxisomes and impaired MCS/transporters by quantitative MS-based proteome studies.
  • Computation of an MCS interaction and cellular response network integrating the results obtained and published data (icw LG).
  • Computational integration of proteomics, lipidomics and metabolomics data from peroxisome-deficient cells and cells with non-functional MCS/transporters (icw LG).

Location: Freiburg, Germany
Supervisor: Prof. Dr. B. Warscheid (bettina.warscheidATbiologie.uni-freiburg.de)
Co-supervisors: Prof. Dr. V. Martins dos Santos, Lifeglimmer GmbH & Prof. Dr. M. Schuldiner, Weizmann Institute of Science
Planned secondments: University of Groningen, Weizmann Institute of Science. Lifeglimmer GmbH
Required subject specific skills and expertise: a master’s degree in biology, biochemistry or a related discipline. Experience in proteomics/mass spectrometry and a strong interest in bioinformatics are advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR9

Reference: ESR10-DE

Are you fascinated by biology and bioinformatics in a dynamic company setting? A PhD position is available at the Data Analytics company Lifeglimmer GmbH, Berlin, Germany.
Project title: Model-driven discovery and network modelling of peroxisomal interactions
Description:

  • To identify novel transporter proteins using whole genome searches, analysis of protein sequences of newly identified proteins by bioinformatics tools, including structure prediction.
  • Carry out bioinformatics analysis for domains/transmembrane regions rational design of proteins in yeast
  • Define flux distributions and phenotypes in altered growth/beta oxidation capacity through metabolomics and modelling (mammalian/human cells)
  • Unravel peroxisomal crosstalk by systems biology approaches
  • Semantically integrate heterogeneous datasets

Location: Berlin, Germany
Supervisor: Prof. Dr. V. Martins dos Santos (vdsATlifeglimmer.com)
Co-supervisors: Prof. Dr. B.M.  Bakker, University Medical Center Groningen & Prof. Dr. B. Warscheid, University of Freiburg
Planned secondments: KU LEUVEN, University Medical Center Groningen, University of Freiburg
Required subject specific skills and expertise:

  • Master degree or equivalent in Computational Biology, Bioinformatics,  Systems Biology, Bio(techno)logy or related field;
  • Solid knowledge of bioinformatics;
  • Basic knowledge in (genome-scale) metabolic modelling;
  • Good understanding of molecular biology/biochemistry;
  • Experience in pathway analyses and/or omics data integration would be of advantage;
  • Experience in analysing omics data as well as interpretation of their biological implications;
  • Basic knowledge of common methods of functional genomics;
  • Experience in discussing results of computational analyses in collaboration with wet-lab scientists;
  • Good programming skills in R (Matlab) and experience in at least one other programming language (Python, Perl, Java, C++);
  • Knowledge of semantic web technologies (XML, RDF, SPARQL) would be an asset;
  • A pragmatic and solution oriented working style;
  • Enthusiasm for communicating with various project partners;

Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR10

Reference: ESR11-BE

A PhD position is available at KU Leuven, Department of Cellular and Molecular Medicine, Leuven, Belgium
Project title: Peroxisomal redox metabolism in health and disease
Description:

  • To identify proteins that have the capacity to channel H2Oacross the peroxisomal membrane using CRISPR-based approaches in combination with a cell line in which the production of peroxisomal H2Ocan be carefully controlled;
  • To study the role of the identified H2Otransporters in signalling pathways and cellular responses to H2O2-induced oxidative insults;
  • To unveil the role of peroxisome membrane contact site (MCS) in interorganellar redox communication (use of synthetic tethers and cell lines in which MCS proteins that will be identified in WP1 are inactivated by CRISPRn);
  • To assess how altered expressions of known and newly-identified peroxisomal transporters modulate redox state in different cell compartments.

Location: Leuven, Belgium
Supervisor: Prof. Dr. M. Fransen (marc.fransenATkuleuven.be)
Co-supervisors: Prof. Dr. E. Gnaiger, Oroboros Instruments GmbH & Prof. dr. H. R. Waterham, Academic Medical Centre at the University of Amsterdam
Planned secondments: The Hospital for Sick Children, Academic Medical Centre at the University of Amsterdam, Oroboros Instruments GmbH
Required subject specific skills and expertise: a master’s degree in life sciences (e.g., biomedical sciences, molecular cell biology, biochemistry, or biophysics). Experience in mammalian cell culture and fluorescence microscopy is advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR11

Reference: ESR12-BE

A PhD position is available at KU Leuven, Department of Cellular and Molecular Medicine, Leuven, Belgium

Project title: Physiological targets of peroxisome-derived hydrogen peroxide
Description:

  • To inventorize protein thiol targets of peroxisome-derived H2O2 in human cells;
  • To study the role of peroxisome-derived H2Oin retrograde signalling;
  • To assess how acute and chronic doses of peroxisome-derived H2Oaffect mitochondrial morphology (e.g. shape, number, size, positioning), function (e.g. respiration), and interorganellar contact sites;
  • To evaluate peroxisome and mitochondrial abundance and morphology in mammalian cells using artificial intelligence-based image analysis methods;
  • To examine how changes in peroxisome abundance and function alter the cellular protein thiol oxidation profile;
  • To validate the in cellulo findings in more disease-related cell types and tissues from control mice and mice suffering from peroxisomal deficiencies.

Location: Leuven, Belgium

Supervisor: Prof Dr. M. Fransen (marc.fransenATkuleuven.be)
Co-supervisors: Prof. Dr. E. Gnaiger, Oroboros Instruments GmbH & Prof. Dr. M. Schrader, University of Exeter
Planned secondments: Oroboros Instruments GmbH, University of Exeter, Lambert Instruments
Required subject specific skills and expertise: a master’s degree in life sciences (e.g., biomedical sciences, molecular cell biology, biochemistry, or biophysics). Experience in mammalian cell culture and fluorescence microscopy is advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR12

Reference: ESR13-IL

A PhD Position is available at Weizmann Institute of Science, Department of Molecular Genetics, Rehovot, Israel
Project title: Peroxisome-mitochondria and peroxisome-LD contact sites in yeast
Description:

  • Identifying novel peroxisome contact site proteins using S. cerevisiae strain collections (libraries) and high content fluorescence microscopy screens. Yeast libraries will be constructed in which each strain will contain a contact site reporter together with one mCherry-tagged yeast protein which is over expressed, covering the entire yeast proteome. The screen will identify which overexpressions affect peroxisome contacts (expansion, increased number per cell etc.) and which proteins reside in the contacts.
  • Identifying peroxisome- mitochondria and peroxisome-lipid droplets (LDs) contact site resident proteins using BioID2 followed by a quantitative proteomics in S. cerevisiae.
  • Discovering the physiological roles of selected, promising peroxisome–LD and/or peroxisome-mitochondrial contact site in S. cerevisiae. Various methods will be used to examine the potential role of these contacts in peroxisome fission, inheritance, ability to perform β-oxidation of fatty acids, detoxification of H2O2 and more.

Location: Rehovot, Israel
Supervisor: Prof. Dr. M. Schuldiner (maya.schuldinerATweizmann.ac.il)
Co-supervisors: Prof. Dr. B. Warscheid, University of Freiburg & Dr. M. Danielsen, MS-Omics ApS
Planned secondments: University of Freiburg, University of Groningen, MS-Omics ApS
Required subject specific skills and expertise: 

  • A master’s degree in molecular life sciences or related fields.
  • Experience in yeast research mandatory.
  • Experience with high content screening approaches mandatory
  • Experience with handling big data sets mandatory
  • Experience with fluorescent microscopes is an advantage
  • Experience in working with both academia and industry is an advantage

Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR13

Reference: ESR14-UK

A PhD Position is available at the University of Exeter, Department of Biosciences, Exeter, United Kingdom
Project title: The peroxisome-ER membrane contact site (MCS) in humans
Description:

  • To identify and characterise interaction partners and regulators of the novel peroxisome-ER MCS mediated by the ACBD4/5-VAPA/B tether in humans (bioinformatics, pull down assays, BioID2, mass spectrometry, molecular cell biology);
  • To develop and establish models and techniques for the analysis of lipid transfer between the ER and peroxisomes;
  • To validate selected candidates (CRISPR, fluorescence and quantitative electron microscopy, cell fractionation) and to reveal their physiological role in health and disease conditions;
  • To examine if mutations in MCS genes can be linked to a collection of fibroblasts obtained from patients with peroxisomal disorders whose genetic bases have not yet been identified.

Location: Exeter, United Kingdom
Supervisor: Prof. Dr. M. Schrader (m.schraderATexeter.ac.uk)
Co-supervisors: Dr. M. Danielsen, MS-Omics ApS & Dr. P. Kim, The Hospital for Sick Children
Planned secondments: The Hospital for Sick Children, Academic Medical Centre at the University of Amsterdam, MS-Omics ApS
Required subject specific skills and expertise: a master’s degree in life sciences/biology (molecular cell biology, biochemistry). Experience in mammalian cell culture, fluorescence microscopy and protein biochemistry is advantageous but not mandatory.
Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR14

Reference: ESR15-UK

A PhD Position is available at the University of Exeter, Department of Biosciences, Exeter, United Kingdom
Project title: Modulating human organelle contacts to improve cell performance
Description:

  • To generate fluorescence-based reporter systems to analyse peroxisome-organelle (mitochondria, ER) interactions;
  • To determine physiological conditions which modulate organelle contacts, e.g. peroxisome-mitochondria interaction during oxidative stress.
  • To perform compound screening to identify modulators of organelle contacts (supported by automated image analysis, LI).
  • To analyse the impact of modulating organelle contacts on cell performance using cell models for oxidative stress and organelle dysfunction (measure organelle/mitochondrial parameters (Seahorse technology, lipid metabolism etc.).

Location: Exeter, United Kingdom
Supervisor: Prof. Dr. M. Schrader (m.schraderATexeter.ac.uk)
Co-supervisors: Dr. D. Hoepfner, Novartis Pharma AG & Prof. Dr. M. Fransen, KU Leuven
Planned secondments: KU Leuven, Novartis Pharma AG, MS-Omics ApS
Required subject specific skills and expertise: a master’s degree in life sciences/biology (molecular cell biology, biochemistry). Experience in mammalian cell culture, fluorescence microscopy, molecular cloning and protein biochemistry is advantageous but not mandatory.

Application deadline: 28/10/2018
Starting date:  April 1, 2019

Application form ESR15

1,842 total views, 6 views today